1. National Institute of Biological Sciences, Zhongguancun Life Science Park, Beijing 102206, China
2. College of Life Sciences, Beijing Normal University, Beijing 100875, China
3. Peking-Yale Joint Center for Plant Molecular Genetics and Agro-Biotechnology, National Laboratory of Protein Engineering and Plant Genetic Engineering, College of Life Sciences, Peking University, Beijing 100871, China
4. Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut 06520-8104
5. State Key Laboratory of Plant Cell and Chromosome Engineering, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing 100101, People's Republic of China