1. Department of Computational and Systems Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom
2. Department of Metabolic Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom
3. Department of Cell and Developmental Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom
4. European Bioinformatics Institute, Hinxton, Cambridge CB10 1SD, United Kingdom
5. Cold Spring Harbor Laboratory, Cold Spring Harbor, New York 11724
6. New York University, School of Medicine, New York, New York 10016