1. Division of Applied Biosciences, Graduate School of Agriculture, Kyoto University, Sakyo-ku, Kyoto 606-8502, Japan
2. Division of Applied Life Sciences, Graduate School of Agriculture, Kyoto University, Sakyo-ku, Kyoto 606-8502, Japan
3. CREST, Japan Science and Technology Agency, 5, Chiyoda-ku, Tokyo 102-0075, Japan