1. Frontier Research Institute for Interdisciplinary Sciences, Tohoku University, 980-8578 Sendai, Japan
2. Department of Environmental Life Sciences, Graduate School of Life Sciences, Tohoku University, 980-8577 Sendai, Japan
3. Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency, 332-0012 Saitama, Japan
4. Department of Applied Plant Sciences, Graduate School of Agricultural Sciences, Tohoku University, 981-8555 Sendai, Japan