1. Machine Learning and Computational Biology Research Group, Max Planck Institute for Intelligent Systems and Max Planck Institute for Developmental Biology, 72076 Tübingen, Germany
2. Zentrum für Bioinformatik, Eberhard Karls Universität, 72074 Tübingen, Germany
3. Department for Biosystems Science and Engineering, ETH Zürich, 4058 Basel, Switzerland
4. Swiss Institute of Bioinformatics, 4058 Basel, Switzerland
5. Department of Molecular Biology, Max Planck Institute for Developmental Biology, 72076 Tübingen, Germany
6. Department of Empirical Inference, Max Planck Institute for Intelligent Systems, 72076 Tübingen, Germany