1. Department of Plant Biology, University of Minnesota–Twin Cities, St. Paul, Minnesota 55108
2. The Sainsbury Laboratory, John Innes Centre, Colney, Norwich NR4 7UH, United Kingdom
3. Department of Plant–Microbe Interactions, Max-Planck-Institute for Plant Breeding Research, Carl-von-Linné-Weg 10, D-50829 Cologne, Germany