1. Integrative Cell Biology Laboratory, School of Biological and Biomedical Sciences, University of Durham, South Road, Durham DH1 3LE, United Kingdom
2. Max-Delbrück Laboratorium, Carl von Linne Weg 10, D-50829 Köln, Germany
3. Plant Science Division, University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5RD, United Kingdom