1. Graduate School of Biostudies, Kyoto University, Kyoto 606-8502, Japan
2. Division of Biological Science, Graduate School of Science, Nagoya University, Chikusa, Nagoya 464-8602, Japan
3. Center for Gene Research, Nagoya University, Chikusa, Nagoya 464-8602, Japan
4. Graduate School of Science, Kobe University, Kobe 657-8501, Japan