1. U.S. Department of Agriculure–Agricultural Research Service, Plant, Soil, and Nutrition Laboratory, Cornell University, Ithaca, New York 14853
2. Department of Plant Biology, Cornell University, Ithaca, New York 14853
3. Boyce Thompson Institute for Plant Research, Cornell University, Ithaca, New York 14853
4. U.S. Department of Agriculure–Agricultural Research Service, Plant Science Research Unit, University of Minnesota, St. Paul, Minnesota 55108
5. Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York 14853