1. Department of Forest Genetics and Plant Physiology, Umeå Plant Science Centre, Swedish University of Agricultural Sciences, SE-901 83 Umeå, Sweden
2. Department of Biology and Institute for Genome Sciences & Policy, Center for Systems Biology, Duke University, Durham, North Carolina 27708, USA
3. Department of Plant Physiology, Umeå Plant Science Centre, Umeå University, SE-901 87 Umeå, Sweden