1. Department of Biological Science, Graduate School of Science, Hiroshima University, Kagamiyama, Higashi-Hiroshima 739-8526, Japan
2. RIKEN Plant Science Center, Tsurumi, Yokohama 230-0045, Japan
3. Department of Biological Sciences, Graduate School of Sciences, University of Tokyo, Hongo, Bunkyo-ku, Tokyo 113-0033, Japan