1. Joint International Research Laboratory of Metabolic & Developmental Sciences, State Key Laboratory of Hybrid Rice, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai 200240, China
2. Department of Life Science and Environmental Biochemistry, Pusan National University, Miryang 50463, Republic of Korea
3. School of Biosciences, University of Melbourne, Parkville, Victoria 3010, Australia
4. Department for Plant and Environmental Sciences, University of Copenhagen, 1871, Frederiksberg C, Denmark
5. Copenhagen Plant Science Center, University of Copenhagen, 1871, Frederiksberg C, Denmark