1. Department of Crop Genetics, John Innes Centre, Norwich NR4 7UH, United Kingdom
2. CSIRO Agriculture and Food, Canberra, ACT 2601, Australia
3. Plant Breeding Institute, School of Life and Environmental Sciences, University of Sydney, Cobbitty, NSW 2570, Australia
4. John Bingham Laboratory, National Institute of Agricultural Botany, Cambridge CB3 0LE, United Kingdom