1. Graduate School of Agricultural and Life Sciences, University of Tokyo, Bunkyo-ku, Tokyo 113-8657, Japan
2. Division of Molecular and Life Sciences, Pohang University of Science and Technology, Pohang 790-784, Republic of Korea
3. National Institute of Agrobiological Sciences, Kannondai, Tsukuba 305-0856, Japan
4. Graduate School of Science, University of Tokyo, Hongo, Tokyo 113-0033, Japan