1. Department of Horticulture, University of Wisconsin-Madison, Madison, Wisconsin 53706
2. Department of Plant Biology, Michigan State University, East Lansing, Michigan 48824
3. Department of Horticulture, Michigan State University, East Lansing, Michigan 48824
4. State Key Laboratory for Crop Genetics and Germplasm Enhancement, Nanjing Agriculture University, Nanjing, Jiangsu 210095, China
5. Jiangsu Collaborative Innovation Center for Modern Crop Production, Nanjing Agriculture University, Nanjing, Jiangsu 210095, China