1. Plant Sciences Division, School of Life Sciences, University of Dundee, Dundee DD2 5DA, United Kingdom
2. Information and Computational Sciences, The James Hutton Institute, Dundee DD2 5DA, United Kingdom
3. Institute of Molecular, Cell, and Systems Biology, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow G12 8QQ, United Kingdom
4. Cell and Molecular Sciences, The James Hutton Institute, Dundee DD2 5DA, United Kingdom
5. Department of Biosciences, Durham University, Durham DH1 3LE, United Kingdom