1. Center of Pear Engineering Technology Research, State Key Laboratory of Crop Genetics and Germplasm Enhancement, College of Horticulture, Nanjing Agricultural University, Nanjing 210095, China
2. College of Horticulture, Fujian Agriculture and Forestry University, Fuzhou 350002, China
3. School of Biosciences, Cardiff University, Museum Avenue, Cardiff CF10 3AX, United Kingdom