1. PlantLab, Institute of Life Sciences, Scuola Superiore Sant’Anna, Pisa 56127, Italy
2. Department of Biology, University of Pisa, Pisa 56124, Italy
3. Max Planck Institute of Molecular Plant Physiology, Potsdam-Golm 14476, Germany
4. Institute of Agricultural Sciences, ETH Zurich, CH-8092 Zurich, Switzerland