1. Environmental Sciences Division, Oak Ridge National Laboratory, Oak Ridge, TN 37831-6422 wullschlegsd@ornl.gov
2. Umeå Plant Science Center, Department of Plant Physiology, Umeå University, SE-901 87 Umeå, Sweden
3. School of Biological Sciences, University of Southampton, Bassett Crescent East, Southampton S016 7PX, UK