1. Department of Biology, University of Saskatchewan, Saskatoon, Saskatchewan S7N 5E2, Canada
2. Laboratory of Cell Biology, College of Life Science and Technology, Huazhong Agricultural University, Wuhan, Hubei 430070, China
3. Saskatoon Research and Development Centre, Agriculture and Agri-Food Canada, Saskatoon, Saskatchewan S7N 0X2, Canada
4. Key Laboratory of Biology and Genetic Improvement of Oil Crops, Ministry of Agriculture, Oil Crops Research Institute, Chinese Academy of Agricultural Sciences, Wuhan, Hubei 430062, China
5. National Research Council Canada, Saskatoon, Saskatchewan S7N 0W9, Canada
6. Global Institute for Food Security, University of Saskatchewan, Saskatoon, Saskatchewan S7N 0W9, Canada
7. Department of Botany and Plant Pathology, Purdue University, West Lafayette, Indiana 47907
8. Laboratoire Reproduction et Développement des Plantes, Université de Lyon, École normale supérieure de Lyon, Université Claude Bernard Lyon 1, CNRS, INRA, Lyon 69342, France