1. Department of Plant Science, University of Adelaide, Waite Campus, Glen Osmond, South Australia 5064, Australia
2. School of Botany, University of Melbourne, Parkville, Victoria 3052, Australia
3. Department of Cell Biology, John Innes Centre, Norwich Research Park, Colney, Norwich NR4 7UH, United Kingdom
4. Sainsbury Laboratory, John Innes Centre, Norwich Research Park, Colney, Norwich NR4 7UH, United Kingdom