1. Laboratory of Plant Biotechnology, University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
2. Department of Applied Chemistry, Science University of Tokyo, Kagurazaka, Shinjuku-ku, Tokyo 162-8601, Japan
3. Laboratory of Plant Molecular Physiology, University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
4. Core Research for Evolutional Science and Technology, Japan Science and Technology Corporation, 332-0012 Saitama, Japan