1. Department of Agronomy, Iowa State University, Ames, Iowa 50011
2. Graduate School of Agricultural and Life Sciences, University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo 113-8657, Japan
3. Center for Plant Genomics, Iowa State University, Ames, Iowa 50011
4. Department of Zoology and Genetics, Iowa State University, Ames, Iowa 50011