1. Graduate School of Bioscience and Biotechnology, Tokyo Institute of Technology, Nagatsuta, Midori-ku, Yokohama 226-8501, Japan
2. Michigan State University–Department of Energy Plant Research Laboratory, Michigan State University, East Lansing, Michigan 48824
3. Department of Plant Biology, Michigan State University, East Lansing, Michigan 48824
4. Boyce Thompson Institute for Plant Research, Ithaca, New York 14853
5. Kazusa DNA Research Institute, Kisarazu, Chiba 292-0812, Japan