1. Experimental Farm/Plant Genetics Laboratory, National Institute of Genetics, Yata1111, Mishima, Shizuoka 411-8540, Japan
2. Department of Molecular Genetics, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki 305-8602, Japan
3. Department of Life Science, Graduate University for Advanced Studies, Yata1111, Mishima, Shizuoka 411-8540, Japan