1. Department of Physiology, Keio University School of Medicine, Tokyo, Japan
2. Faculty of Life and Environmental Sciences, Graduate Faculty of Interdisciplinary Research, University of Yamanashi, Yamanashi, Japan
3. Center for Genomic and Regenerative Medicine, Juntendo University School of Medicine, Tokyo, Japan
4. Department of Health Sciences, Faculty of Medicine, University of Yamanashi, Yamanashi, Japan
5. Department of Dermatology, Keio University School of Medicine, Tokyo, Japan
6. Present address: Dermatology Branch, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, MD 20892, USA
7. Present address: Department of Dermatology, Kyorin University School of Medicine, Tokyo, Japan
8. Department of Epigenetic Medicine, Faculty of Medicine, University of Yamanashi, Yamanashi, Japan