1. Department of Biological Sciences, Dartmouth College, Hanover NH 03755
2. Center for Molecular Medicine, Molecular Cancer Research, University Medical Center Utrecht, 3584 CG Utrecht, The Netherlands, and
3. King Abdullah University of Science and Technology (KAUST), Biological and Environmental Science and Engineering Division (BESE), KAUST Environmental Epigenetics Program (KEEP), Thuwal 23955-6900, Saudi Arabia