1. Triticeae Research Institute, Sichuan Agricultural University at Chengdu, Wenjiang, Sichuan 611130, People’s Republic of China
2. Institute of Ecological Forestry, Sichuan Agricultural University at Chengdu, Wenjiang, Sichuan 611130, People’s Republic of China
3. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810001, People’s Republic of China
4. INRA University Blaise Pascal, Joint Research Unit 1095 Genetics Diversity and Ecophysiology of Cereals, Clermont-Ferrand 63039, France
5. Department of Biology and Microbiology, South Dakota State University, Brookings, South Dakota 57007