1. Departamento de Bioquímica, Genética e Inmunología, Facultad de Biología, Universidad de Vigo, Vigo 36310, Spain
2. The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian EH25 9RG, Scotland, United Kingdom
3. Medical Research Council (MRC) Human Genetics Unit (HGU) at the MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh EH4 2XU, Scotland, United Kingdom
4. Institute of Evolutionary Biology, University of Edinburgh, Edinburgh EH9 3FL, Scotland, United Kingdom