1. CBGP, INRA, CIRAD, IRD, Montpellier SupAgro, Univ Montpellier, 34988 Montferrier-sur-Lez Cedex, France
2. Institut de Biologie Computationnelle, Univ Montpellier, 34095 Montpellier Cedex, France
3. ESE, Ecology and Ecosystem Health, INRA, Agrocampus Ouest, 35042 Rennes, Cedex, France