1. Department of Microbiology and Institute of Cancer Research, Columbia University, New York, New York 10032
2. Department of Microbiology, University College Cork, Cork, Ireland
3. Center for Bioimage Informatics and Department of Biomedical Engineering, Carnegie Mellon University, Pittsburgh, Pennsylvania 15213
4. Lane Center for Computational Biology and Department of Machine Learning, Carnegie Mellon University, Pittsburgh, Pennsylvania 15213
5. External Fellow, Freiburg Institute for Advanced Studies, University of Freiburg, 79104 Freiburg, Germany and
6. Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania 15213