1. A*STAR Institute of Molecular and Cell Biology, Singapore 138673, Singapore
2. Lee Kong Chian School of Medicine, Nanyang Technological University, Singapore 639798, Singapore
3. The Living Systems Institute, University of Exeter, EX4 4QD UK
4. Department of Biomedical Science, University of Sheffield, S10 2TN, UK
5. Temasek Lifesciences Laboratory, Singapore 117604, Singapore