1. Chinese Academy of Sciences Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Laboratory for Marine Ecology and Environmental Science, Qingdao National Laboratory for Marine Science and Technology, 266071, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. College of Fisheries, Ocean University of China, Qingdao, 266001 China