1. Department of Fish Ecology and Evolution, Centre of Ecology, Evolution and Biogeochemistry, EAWAG Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland
2. Division of Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Switzerland
3. Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany