1. Department of Forest and Conservation Sciences, The University of British Columbia, Vancouver, BC, Canada V6T 1Z4
2. Department of Wood Science, The University of British Columbia, Vancouver, BC, Canada V6T 1Z4
3. Michael Smith Laboratory, The University of British Columbia, Vancouver, BC, Canada V6T 1Z4
4. Natural Resources Canada, Laurentian Forestry Centre, Québec, QC, Canada G1V 4C7