1. a Department of Biology , University of Nevada–Reno , Mail Stop 314, Reno , Nevada , 89557 , USA
2. b U.S. Fish and Wildlife Service , Lahontan National Fish Hatchery Complex , 1340 Financial Boulevard, Suite 234, Reno , Nevada , 89502 , USA
3. c Department of Watershed Sciences , Utah State University , 5210 Old Main, Logan , Utah , 84321 , USA