1. a U.S. Geological Survey , 2795 Anderson Avenue, Suite 106, Klamath Falls , Oregon , 97603 , USA
2. b U.S. Geological Survey, Washington Cooperative Fish and Wildlife Research Unit School of Aquatic and Fishery Sciences, University of Washington, Box 355020 , Seattle , Washington , 98195 , USA
3. c U.S. Geological Survey , Oregon Water Science Center, 2130 SW 5th Avenue , Portland , Oregon , 97201 , USA