1. Terrestrial Ecology Department; Netherlands Institute of Ecology (NIOO-KNAW); Droevendaalsesteeg 10 6708 PB Wageningen the Netherlands
2. Ecology & Evolutionary Biology; Princeton University; Princeton NJ 08544 USA
3. Ecology & Evolutionary Biology; University of California; Los Angeles CA 90095 USA
4. Institute of the Environment and Sustainability; University of California; Los Angeles CA 90095 USA