1. State Key Laboratory of Estuarine and Coastal Research; East China Normal University; Shanghai 200062 China
2. Laboratory of Protozoology; Institute of Evolution & Marine Biodiversity; Ocean University of China; Qingdao 266003 China
3. College of Fisheries and Life Science; Shanghai Ocean University; Shanghai 201302 China
4. College of Life Sciences; University of Chinese Academy of Sciences; Beijing 100049 China
5. Zoology Department; College of Science; King Saud University; Riyadh 11451 Saudi Arabia