1. Guangdong Provincial Key Laboratory for Plant Epigenetics, Longhua Bioindustry and Innovation Research Institute, College of Life Sciences and Oceanography Shenzhen University Shenzhen Guangdong 518060 China
2. Hefei National Laboratory for Physical Sciences at the Microscale, CAS Center for Excellence in Molecular Plant Sciences, School of Life Sciences Division of Life Sciences and Medicine, University of Science and Technology of China Hefei 230027 China
3. Department of Botany and Plant Sciences, Institute for Integrative Genome Biology University of California Riverside CA 92521 USA
4. State Key Laboratory of Genetic Engineering and Ministry of Education Key Laboratory of Biodiversity Sciences and Ecological Engineering, Institute of Plant Biology, School of Life Sciences Fudan University Shanghai 200438 China