1. Department of Organismic and Evolutionary Biology; Harvard University; 16 Divinity Ave Cambridge MA 02138-2020 USA
2. SOA Key Laboratory for Polar Science; Polar Research Institute of China; Shanghai 200136 China
3. Department of Marine Chemistry and Geochemistry; Woods Hole Oceanographic Institution; 266 Woods Hole Rd., MS 25 Woods Hole MA 02540 USA
4. College of Life Sciences; Zhejiang University; 388 Yuhangtang Road Hangzhou 310058 China