1. School of Life Sciences Lanzhou University Lanzhou China
2. Department of Geosciences and Geography University of Helsinki Helsinki Finland
3. Department of Ecology and Evolution University of Lausanne Lausanne Switzerland
4. College of Ecology Lanzhou University Lanzhou China
5. Institute of Earth Surface Dynamics University of Lausanne Lausanne Switzerland
6. INRAE Université de Lyon, Université Lyon 1, CNRS, vetAgroSup, UMR 1418 INRAE, UMR 5557 CNRS, Ecologie Microbienne LEM Villeurbanne France