1. Graduate School of Agriculture Kyoto University Kyoto 606‐8502 Japan
2. Department of Biological Sciences Macquarie University Sydney NSW 2109 Australia
3. Division of Plant Sciences Research School of Biology The Australian National University Canberra ACT 0200 Australia
4. Graduate School of Life Sciences Tohoku University Aoba Sendai 980‑8578 Japan
5. Institute of Agricultural and Environmental Sciences Estonian University of Life Sciences Tartu 51014 Estonia
6. Plant Sciences (IBG‐2) Forschungszentrum Jülich GmbH D‐52425 Jülich Germany