1. Australian Wetlands, Rivers and Landscapes Centre; The University of New South Wales; Sydney NSW 2052 Australia
2. Evolution & Ecology Research Centre; The University of New South Wales; Sydney NSW 2052 Australia
3. School of Mathematics and Statistics; The University of New South Wales; Sydney NSW 2052 Australia
4. Australian Museum; Sydney NSW 2010 Australia
5. School of the Environment; University of Technology Sydney; Sydney NSW 2007 Australia