1. Institute of Biodiversity, Animal Health & Comparative Medicine; University of Glasgow; Glasgow G12 8QQ U.K.
2. Scottish Natural Heritage; Caspian House, Mariner Court,; Clydebank, G81 2NR U.K.
3. Scottish Centre for Ecology & the Natural Environment, IBAHCM, Rowardennan, Loch Lomond G63 0AW; U.K.