1. Centre for Geographic Studies, University of Lisbon; Alameda da Universidade; 1600-214 Lisbon; Portugal
2. Department of Ecology and Evolutionary Biology; University of Tennessee; Knoxville; TN; 37917; U.S.A.
3. Department of Biology; University of Florence; via Romana 17; 50125 Florence; Italy
4. School of Aquatic and Fishery Sciences; University of Washington; Seattle; WA; 98195; U.S.A.