1. College of Grassland Science Beijing Forestry University Beijing China
2. Key Laboratory of Alpine Ecology Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing China
3. University of Chinese Academy of Sciences Beijing China
4. CAS Center for Excellence in Tibetan Plateau Earth Sciences Beijing China
5. Northwest Institute of Plateau Biology Chinese Academy of Sciences Xining China
6. Qinghai Provincial Key Laboratory of Restoration Ecology of Cold Area Xining China
7. Earth and Environmental Sciences Area Lawrence Berkeley National Laboratory Berkeley CA USA
8. Biosciences Area Lawrence Berkeley National Laboratory Berkeley CA USA
9. The University of Chicago Medicine and Biological Sciences Division Chicago IL USA