1. Institute of Biology and Environmental Sciences; Functional Ecology of Plants; Carl-von-Ossietzky University Oldenburg; PO Box 2503 26111 Oldenburg Germany
2. Institute of Biology and Environmental Sciences; Biodiversity and Evolution of Plants; Carl-von-Ossietzky University Oldenburg; PO Box 2503 26111 Oldenburg Germany
3. Smithsonian Tropical Research Institute; Apartado Postal 0843-03092 Panamá República de Panamá