1. Department of Biology; University of Iowa; Iowa City IA 52242 USA
2. Department of Systematic and Evolutionary Botany; University of Zurich; Zurich CH-8008 Switzerland
3. Swiss Institute of Bioinformatics; Quartier Sorge-Batiment Genopode; Lausanne Switzerland
4. MTA ELTE-MTM Ecology Research Group; ELTE; Biological Institute; Budapest Hungary