1. AIMS@JCU; Australian Institute of Marine Science; School of Marine and Tropical Biology; James Cook University; Townsville Qld 4811 Australia
2. Centre for Sustainable Tropical Fisheries and Aquaculture; James Cook University; Townsville Qld 4811 Australia
3. Department of Environment and Agriculture; Curtin University; Perth WA 6845 Australia
4. ARC Centre of Excellence for Coral Reef Studies; James Cook University; Townsville Qld 4811 Australia
5. Australian Institute of Marine Science; PMB 3; Townsville MC Townsville Qld 4810 Australia
6. School of Marine and Tropical Biology; James Cook University; Townsville QLD 4811 Australia